这一次,破坏王拉尔夫(约翰·C·赖利 John C. Reilly 配音)和云妮洛普(萨拉·丝沃曼 Sarah Silverman 配音)又闯祸了,导致了云妮洛普所在的游戏《甜蜜冲刺》的游戏机方向盘遭到了损坏。这是一台非常古老的街机,在整个互联网中,只有一只备用方向盘正在销售,然而它的售价实在是非常昂贵,老板决定放弃这台机器,将它解体变卖。这也就意味着,《甜蜜冲刺》里的角色们将永远失去他们的家园。 为了避免这一悲剧的发生,拉尔夫和云妮洛普决定亲自到互联网里去走一遭,买回方向盘。一路上,他们惹了无数的笑料,犯了无数令人啼笑皆非的错误。为了赚钱,拉尔夫不惜亲自出马成为视频网站的网红赚取点击量变现,而云妮洛普则在另一款赛车游戏《狂野飙车》里找到了真正的自我。
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